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The first evidence that primary cilia might activate intracellular pathways upon mechanical bending came from landmark studies that demonstrated that bending of primary cilia either by applying an apical membrane flow on renal epithelial cells (MDCK) grown polarized, or by pulling on primary cilia using a micropipette, was sufficient to elicit an intracellular calcium release signal from intracellular ER storage sites mediated by IP3 receptors (68)